FORM 4 Check this box if no longer subject to Section 16. Form 4 or Form 5 obligations may continue. See Instruction 1(b).

UNITED STATES SECURITIES AND EXCHANGE COMMISSION Washington, D.C. 20549 STATEMENT OF CHANGES IN BENEFICIAL OWNERSHIP OF SECURITIES Filed pursuant to Section 16(a) of the Securities Exchange Act of 1934, Section 17(a) of the Public Utility Holding Company Act of 1935 or Section 30(h) of the Investment Company Act of 1940

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(Print or Type Responses)

1. Name and Address of Reporting Person * WHITE TONY L   2. Issuer Name and Ticker or Trading Symbol APPLERA CORP [ABI/CRA] 5. Relationship of Reporting Person(s) to Issuer (Check all applicable) __X__ Director _____ 10% Owner __X__ Officer (give title below) _____ Other (specify below) Chairman, President and CEO (Last) (First) (Middle) APPLERA CORPORATION, 301 MERRITT 7 3. Date of Earliest Transaction (Month/Day/Year) 06/30/2006 (Street) NORWALK, CT 06851-1070 4. If Amendment, Date Original Filed(Month/Day/Year) 6. Individual or Joint/Group Filing(Check Applicable Line) _X_ Form filed by One Reporting Person ___ Form filed by More than One Reporting Person (City) (State) (Zip) Table I - Non-Derivative Securities Acquired, Disposed of, or Beneficially Owned

1.Title of Security (Instr. 3) 2. Transaction Date (Month/Day/Year) 2A. Deemed Execution Date, if any (Month/Day/Year) 3. Transaction Code (Instr. 8) 4. Securities Acquired (A) or Disposed of (D) (Instr. 3, 4 and 5) 5. Amount of Securities Beneficially Owned Following Reported Transaction(s) (Instr. 3 and 4) 6. Ownership Form: Direct (D) or Indirect (I) (Instr. 4) 7. Nature of Indirect Beneficial Ownership (Instr. 4) Code V Amount (A) or (D) Price Applied Biosystems Group Common Stock 06/30/2006   F   9,834 D $ 32.67 500,948.2486 (1) (2) (3) D   Applied Biosystems Group Common Stock               75,000 I By Elizabeth Ann White Business Interests, LLLP (4) Applied Biosystems Group Common Stock 05/04/2006   G(5) V 15,930 D $ 0 0 (2) I By Tony Lee White 2004 Two Year Grantor Retained Annuity Trust (6) Applied Biosystems Group Common Stock               63,000 (3) I By Tony Lee White 2006 Two Year Grantor Retained Annuity Trust Celera Genomics Group Common Stock 06/30/2006   F   3,278 D $ 13.01 103,154.3648 (1) (3) (7) D   Celera Genomics Group Common Stock               25,000 I By Elizabeth Ann White Business Interests, LLLP (4) Celera Genomics Group Common Stock 05/04/2006   G(5) V 5,310 D $ 0 0 (7) I By Tony Lee White 2004 Two Year Grantor Retained Annuity Trust (6) Celera Genomics Group Common Stock               21,000 (3) I By Tony Lee White 2006 Two Year Grantor Retained Annuity Trust

Reminder: Report on a separate line for each class of securities beneficially owned directly or indirectly.
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Table II - Derivative Securities Acquired, Disposed of, or Beneficially Owned
(e.g., puts, calls, warrants, options, convertible securities)

1. Title of Derivative Security (Instr. 3)

2. Conversion or Exercise Price of Derivative Security

3. Transaction Date (Month/Day/Year)

3A. Deemed Execution Date, if any (Month/Day/Year)

4. Transaction Code (Instr. 8)

5. Number of Derivative Securities Acquired (A) or Disposed of (D) (Instr. 3, 4, and 5)

6. Date Exercisable and Expiration Date (Month/Day/Year)

7. Title and Amount of Underlying Securities (Instr. 3 and 4)

8. Price of Derivative Security (Instr. 5)

9. Number of Derivative Securities Beneficially Owned Following Reported Transaction(s) (Instr. 4)

10. Ownership Form of Derivative Security: Direct (D) or Indirect (I) (Instr. 4)

11. Nature of Indirect Beneficial Ownership (Instr. 4)

Code

V

(A)

(D)

Date Exercisable

Expiration Date

Title

Amount or Number of Shares

Reporting Owners

Reporting Owner Name / Address	Relationships
	Director	10% Owner	Officer	Other
WHITE TONY L APPLERA CORPORATION 301 MERRITT 7 NORWALK, CT 06851-1070	X		Chairman, President and CEO

Signatures

/s/ Thomas P. Livingston, Attorney-in-Fact for Tony L. White		07/05/2006
**Signature of Reporting Person		Date

Explanation of Responses:

*	If the form is filed by more than one reporting person, see Instruction 4(b)(v).
**	Intentional misstatements or omissions of facts constitute Federal Criminal Violations. See 18 U.S.C. 1001 and 15 U.S.C. 78ff(a).
(1)	Includes 471.2535 shares of Applied Biosystems Group Common Stock ("ABI stock") and 1,144.6886 shares of Celera Genomics Group Common Stock ("Celera stock") purchased in March 2006 under the issuer's employee stock purchase plan.
(2)	Reflects the final distribution of 32,550 shares of ABI stock by the Tony Lee White 2004 Two Year Grantor Retained Annuity Trust (the "2004 GRAT") to the reporting person on May 4, 2006. The reporting person previously reported these shares as held indirectly through the 2004 GRAT.
(3)	Reflects the transfer of shares of ABI stock and Celera stock by the reporting person to the Tony Lee White 2006 Two Year Grantor Retained Annuity Trust, of which the reporting person's wife is the sole trustee.
(4)	Elizabeth Ann White Business Interests, LLLP, is a family limited partnership (the "FLP"). The reporting person's adult daughter is the sole general partner of the FLP, and the reporting person's wife and a grantor retained annuity trust established by the reporting person are currently the sole limited partners of the FLP.
(5)	Reflects gift of shares of ABI stock and Celera stock to the reporting person's adult daughter upon the final distribution of the 2004 GRAT.
(6)	The reporting person's wife was the sole trustee of the 2004 GRAT.
(7)	Reflects the final distribution of 10,850 shares of Celera stock by the 2004 GRAT to the reporting person on May 4, 2006. The reporting person previously reported these shares as held indirectly through the 2004 GRAT.

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